分类： 生物学 >> 动物学 提交时间： 2022-09-29 合作期刊: 《古脊椎动物学报》
摘要： Abundant mammalian fossils were uncovered during the field exploration for Nihewan beds at the beginning of the 1980s along Xinyaozi Ravine at Nangaoya Township of Tianzhen County, Shanxi Province in North China. But most equid material was not yet described except that of Equus stenonis. Six forms of Nihewanian equids were confirmed from the Xinyaozi specimens in the present study, five of which were described for the first time. They include four stenonids such as Equus sanmeniensis, E. teilhardi, E. huanghoensis and E. stenonis, and two hipparionines such as Hipparion (Proboscidipparion) sinense and H. (Plesiohiparrion) shanxiense. The diversification of stenonids in the Early Pleistocene was significant in North China with four taxa in Xinyaozi alone. The persistence of Neogene relics such as hipparionines was still present in the Early Pleistocene with two hipparionine taxa in Xinyaozi. Equus sanmeniensis and H. (Proboscidipparion) sinense were two representative equids not only coexisted in the Early Pleistocene but also widely distributed in China. The diversity of equids also implies the diversified vegetation on which they depended. The hypsodont dentitions and well developed cement, as well as completely molarized premolars of Xinyaozi equids indicate their abrasive diet mostly on monocotyledonous and grassland habitats with considerable scales enough to nourish six taxa of equids.
分类： 生物学 >> 动物学 提交时间： 2021-07-21 合作期刊: 《古脊椎动物学报》
摘要： Abundant mammalian fossils were uncovered during the field exploration for Nihewan beds at the beginning of the 1980s along Xinyaozi Ravine at Nangaoya Township of Tianzhen County, Shanxi Province in North China and the studied taxa indicate an age of the early Early Pleistocene. Recent studies on the rhino material not yet described show that there are at least two species of rhinocerotids: Elasmotherium peii and Coelodonta nihowanensis. There might be a third taxon provisionally named as Stephanorhinus cf. S. kirchbergensis due to incompleteness of the specimens. Since its morphometric characters are between S. kirchbergensis and C. nihowanensis, it might be a variety of one of the two species although it is more similar to the former than the latter. In the same way, The rhino specimens from Xiashagou named as Rhinoceros sinensis (?) by Teilhard de Chardin and Piveteau (1930) might be a variety of S. kirchbergensis or C. nihowanensis. The rhinocerotids uncovered so far from the Early Pleistocene deposits in the generalized Nihewan Basin including two certain species and two uncertain ones. The localities yielding E. peii include Xiashagou, Shanshenmiaozhui, Daheigou and Xinyaozi; those yielding C. nihowanensis include Xiashagou, Danangou, Donggutuo, Shanshenmiaozhui and Xinyaozi. R. sinensis (?) appeared only at Xiashagou and Stephanorhinus cf. S. kirchbergensis only at Xinyaozi.
分类： 生物学 >> 动物学 提交时间： 2020-09-15 合作期刊: 《古脊椎动物学报》
摘要： Neogene strata rich in fossil mammals are well exposed in central Nei Mongol, China. Over a dozen mammalian faunas in chronological succession from Early Miocene to Early Pliocene were discerned in this region, and they built a fundamental part of the Neogene land mammal biochronological system in northern China. Tuchengzi was first recognized for producing abundant mammalian fossils during the Sino-Soviet Paleontological Expedition (SSPE) initiated in 1959. However, all unearthed fossils from the SSPE were either large- or middle-sized mammals, and small mammal fossils in the Tuchengzi Fauna were deficient for a long time. The composition and biochronological position of the Tuchengzi Fauna in the Neogene mammalian biochronological system in northern China was not particularly clear. The new fossils here reported are represented by 6 taxa of small mammals. Based on the co-occurrence of Lophocricetus grabaui, Sinocricetus zdanskyi, Prosiphneus licenti, Hansdebruijnia pusilla, and Moschus grandaevus, the age of the Tuchengzi Fauna is constrained to late Late Miocene or Baodean Chinese Land Mammal Age, slighter younger than the Baogeda Ula Fauna and older than the Ertemte Fauna. Judging from the fossil composition, the Tuchengzi Fauna possibly inhabited a forest-steppe mixed habitat during the late Late Miocene.
分类： 生物学 >> 动物学 提交时间： 2020-03-31 合作期刊: 《古脊椎动物学报》
摘要： Many cervid specimens were uncovered during the field exploration for Nihewan beds at the beginning of the 1980s from Taijiaping, Shuichongkou and Dazhuangke localities along Xinyaozi Ravine at Nangaoya Township of Tianzhen County, Shanxi Province in North China. Recent studies on the cervid material identified seven species of Cervidae in total: Muntiacus bohlini, Cervavitus cf. C. huadeensis, Axis shansius, Nipponicervus elegans, Elaphurus davidianus predavidianus, E. bifurcatus from the Early Pleistocene deposits at Taijiaping and Shuichongkou localities, and Cervus (Elaphus) elaphus from uncertain horizon at Dazhuangke. At least the previous six species of cervids were from Nihewan Formation (Nihewanian or equivalent to European middle and late Villafranchian), i.e. the Early Pleistocene, in Sangganhe Basin area.Cervavitus cf. C. huadeensis and A. shansius were survivors from the Late Neogene; M. bohlini,N. elegans, E. davidianus predavidianus and E. bifurcatus are new forms of the Early Pleistocene.If Dazhuangke horizon can be dated as those of Shuichongkou and Taijiaping localities, the appearance of elaphoid cervids could be traced back to the Early Pleistocene, and the evolution of elaphoid antler would start from absence to presence of bez tine. The presence of Elaphurus and Nipponicervus in mainland China and Japanese archipelago implies further that the sea level was dropped down that these cervids could migrate from the mainland to the islands. The abundance of folivorous cervid specimens in the Xinyaozi Ravine area indicates the existence of a certain scale of forested environment in Sangganhe Basin area during the Early Pleistocene.
分类： 生物学 >> 动物学 提交时间： 2019-06-11 合作期刊: 《古脊椎动物学报》
摘要： There are many localities yielding the Pleistocene mammalian faunas in China. It offers excellent material for the study of mammalian evolution, biochronology,paleoecology,paleoenvironment, paleozoogeography, etc. Faunal assemblage characters and taxonomic extinction rates were widely used for determining the faunal ages in biochronology. Faunal binary similarity coefficients sequenced according to Brainerd-Robinson’s rule and antiquity coefficients were further developed methods in biochronology for dating the faunal ages. The faunal binary similarity coefficients are based on the presence or absence of a taxon in a fauna. It is similar to the presence or absence of a character of a species in cladistic analyses for phylogeny, and all faunas have a special ancestor-descendant relationship. The present work is an attempt to find the relationship of the faunas with cladistic methods by selecting three groups of faunas sequenced by faunal binary similarity coefficients according to Brainerd-Robinson’s rule and antiquity coefficients, to compare the results with different methods, and then to estimate the ages of the faunas not yet dated by physical or chemical methods. The estimations are as follow: Gulongshan in Dalian, Liaoning Province: 16–20 ka; Shanchengzi at Benxi, Liaoning Province: 20–30 ka; Hualongdong at Dongzhi Man site, Anhui Province: 150–400 ka; Xinghuashan at Nanzhao Man site, Henan Province: 150–400 ka; Donghe at Luonan Man site, Shaanxi Province: 500–700 ka; Bailongdong at Yunxi Man site, Hubei Province: 500–850 ka; Meipu at Yunxian, Hubei Province: 500–850 ka; Mohui at Tiandong Man site, Guangxi Autonomous Region: 1.2–1.8 Ma; Juyuandong at Liucheng, Guangxi Autonomous Region: 1.2–1.5 Ma; Chutoulang at Chifeng, Nei Mongol Autonomous Region: 1.6–1.9 Ma; Renzidong at Fanchang, Anhui Province: 1.9–2.4 Ma.
分类： 生物学 >> 动物学 提交时间： 2019-06-11 合作期刊: 《古脊椎动物学报》
摘要： There are many localities yielding the Pleistocene mammalian faunas in China. It offers excellent material for the study of mammalian evolution, biochronology,paleoecology,paleoenvironment, paleozoogeography, etc. Faunal assemblage characters and taxonomic extinction rates were widely used for determining the faunal ages in biochronology. Faunal binary similarity coefficients sequenced according to Brainerd-Robinson’s rule and antiquity coefficients were further developed methods in biochronology for dating the faunal ages. The faunal binary similarity coefficients are based on the presence or absence of a taxon in a fauna. It is similar to the presence or absence of a character of a species in cladistic analyses for phylogeny, and all faunas have a special ancestor-descendant relationship. The present work is an attempt to find the relationship of the faunas with cladistic methods by selecting three groups of faunas sequenced by faunal binary similarity coefficients according to Brainerd-Robinson’s rule and antiquity coefficients, to compare the results with different methods, and then to estimate the ages of the faunas not yet dated by physical or chemical methods. The estimations are as follow: Gulongshan in Dalian, Liaoning Province: 16–20 ka; Shanchengzi at Benxi, Liaoning Province: 20–30 ka; Hualongdong at Dongzhi Man site, Anhui Province: 150–400 ka; Xinghuashan at Nanzhao Man site, Henan Province: 150–400 ka; Donghe at Luonan Man site, Shaanxi Province: 500–700 ka; Bailongdong at Yunxi Man site, Hubei Province: 500–850 ka; Meipu at Yunxian, Hubei Province: 500–850 ka; Mohui at Tiandong Man site, Guangxi Autonomous Region: 1.2–1.8 Ma; Juyuandong at Liucheng, Guangxi Autonomous Region: 1.2–1.5 Ma; Chutoulang at Chifeng, Nei Mongol Autonomous Region: 1.6–1.9 Ma; Renzidong at Fanchang, Anhui Province: 1.9–2.4 Ma.
分类： 地球科学 >> 地质学 提交时间： 2018-03-02 合作期刊: 《古脊椎动物学报》
摘要： Moschus grandaevus was firstly uncovered from the Late Miocene deposits at Ertemte, Olan Chorea, Harr Obo and Hua Ba in or around Huade County in the middle part of Nei Mongol and was described by Schlosser in 1924. The excavations by Sino-Soviet Joint Paleontological Team in 1959 and recent excavations since 2013 at Tuchengzi (Tuchetse), another locality at Huade, accumulated many specimens of the musk deer. The morphology and metric studies show that the musk deer specimens from Tuchengzi are the same as those described by Schlosser and can be included into the same species. M. grandaevus ranges from Siberia of Russia to North China, and likely to southern China, in the Late Miocene and Pliocene. The appearance of such folivorous musk deer in the Late Miocene deposits at Tuchengzi indicates that there were forests there during that period. The cladistic analyses show that the fossil species of Moschus are closely related to each other and can be grouped together as Moschini or Moschinae. Micromeryx is closer to Moschus and Cervidae, but the relationship between Micromeryx and Moschus is more complicated than previously considered; nonetheless Hispanomeryx is closer to Bovidae. Schlosser (1924)记述了最初发现于内蒙古化德境内或附近的二登图、敖兰卓蕾、哈尔敖包及华坝的古麝(Moschus grandaevus)。1959年中苏古生物考察队在化德的另一个地点土城子采集到不少古麝化石。2013年以来作者在土城子进行的野外发掘积累了更多的古麝化石标本。土城子标本在形态和测量数据方面和二登图等地点的古麝非常接近，可以归入同一种。根据现有资料归纳，古麝的地理分布范围自俄罗斯的西伯利亚到华北，很可能延伸到华南；其地质时代分布范围为晚中新世–上新世。食叶型古麝在土城子晚中新世地层中的出现说明当时土城子一带有森林环境。支序分析显示麝属(Moschus)化石种相互之间的系统关系非常近，可以归并为麝族(Moschini)或麝亚科(Moschinae); Micromeryx更接近麝属和鹿科，它与麝属之间的关系比以前认为的复杂，而Hispanomeryx更接近牛科。
分类： 生物学 >> 动物学 提交时间： 2017-11-07 合作期刊: 《古脊椎动物学报》
摘要： New specimens from a new locality at Chutoulang in eastern Nei Mongol were identified as Canis chihliensis, Coelodonta nihowanensis, Hipparion (Proboscidipparion) sinense, Equus sanmeniensis, Sus lydekkeri, Muntiacus cf. M. lacustris, Axis shansius, Eucladoceros boulei, Spirocerus cf. S. wongi and Bison palaeosinensis. They enriched the mammalian fauna of Chutoulang to 30 taxa together with the material from Dongliang, Dongcun Beigou and Dongcun Nangou localities. The mammalian fossils from these four localities are all from the same horizon and can be regarded as the same fauna, i.e. Chutoulang fauna. The composition of Chutoulang fauna is the closest to that of Nihewan (s.s.) by binary faunal similarity coefficients. The age of Chutoulang fauna is between those of Nihewan (s.s.) and Juyuandong at Liucheng according to Brainerd-Robinson’s sequence, extinction rates and antiquity coefficients. The numerical age of Chutoulang fauna is estimated between 1.4–1.6 Ma based on those of compared faunas. Carnivora are the most numerous in Chutoulang fauna with 11 taxa, but mostly the small sized ones. Perissodactyla and Artiodactyla make about half of the fauna. They are mostly large sized forms. The presence of numerous browsers or forest dwellers implies the existence of forest or woodland in Chutoulang area during that period. The presence of grazers and openland dwellers indicates the existence of larger area of grassland or steppes than that of woodland or forests. Most members of Chutoulang fauna are temperate habitat dwellers with a few cold-prone forms such as Ochotona and Coelodonta. The climate in Chutoulang area in the Early Pleistocene was thus similar to that of today. Chutoulang fauna is the most northeastern Early Pleistocene fauna in China and it can be recommended as a type site of the Early Pleistocene mammalian fauna in northeastern China.
分类： 地球科学 >> 地质学 提交时间： 2017-08-24 合作期刊: 《古脊椎动物学报》
摘要： Five taxa of Cervidae were identified from the new collection at Tuchengzi locality in Huade, Nei Mongol: Eostyloceros blainvillei, E. triangularis, Euprox sp., Cervavitus huadeensis and C. shanxius, among which the first three taxa were discovered for the first time in Huade area, and the dental specimens of the last two taxa from Tuchengzi locality are also described for the first time. E. blainvillei is a large muntiacine, diagnosed by long, thick and medially curved main beam and a relatively long brow tine emerging directly from the burr. It was originally found in Yushe Basin in Shanxi Province, and reported later from Qaidam Basin in Qinghai Province. Huade is the third area yielding the species. E. triangularis distinguishes from E. blainvillei by its special main beam with triangular cross sections, and it was reported only from Yushe Basin. Its presence in Huade extended its geographic distribution northward to Nei Mongol. Euprox is a transitional form of cervids from permanent antlers to seasonally deciduous ones and it is found in several localities across Eurasia. Huade is the third area in Nei Mongol yielding the taxon after Tung Gur and Siziwang Qi (Siziwang Banner). C. huadeensis is a pliocervine with four tines, and particularly with two distal tines sword-like. It is found at Tuchengzi for the second time, but not elsewhere so far, and it seems to be an endemic taxon. C. shanxius distinguishes from C. novorossiae by absence of Palaeomeryx fold on lower molars. It is widely found in Shanxi, Shaanxi, Gansu and Nei Mongol of northern China. It is often found with numerous specimens in a locality that indicates it lived in large herds. E. blainvillei, E. triangularis and C. shanxius are main members of Yushe I, i.e. Mahui Formation or Baodean. Euprox appeared also mainly in the Late Miocene. The geological age of Tuchengzi locality based on excavated cervids is therefore the Late Miocene. The numerous specimens of C. shanxius indicates forested environment in Huade area in that age. 继1959年中苏古生物联合考察队在内蒙古化德地区发掘采集到大量晚中新世哺乳动物化石后，笔者的课题组于2013~2016年间在化德的土城子地点发掘采集到不少哺乳动物化石。经过修理和研究，鉴定出鹿科化石的5个种类：布氏始柱角鹿(Eostyloceros blainvillei), 三角始柱角鹿(E. triangularis), 真角鹿未定种(Euprox sp.), 化德祖鹿(Cervavitus huadeensis)和山西祖鹿(C. shanxius)。其中前3个种类是在化德地区首次发现，而后两个种的牙齿材料也是在土城子地点首次记述。布氏始柱角鹿是一种较大的麂类，以其鹿角具一个粗长而内弯的主枝及一个直接从角环上伸出的较长的眉枝为特征，以前主要发现于山西的榆社盆地，在青海柴达木盆地也找到过，而化德是出现这个种类的第三个地区。三角始柱角鹿以其主枝横切面呈三角形而与布氏始柱角鹿相区别，以前仅见于榆社盆地，因此新发现的三角始柱角鹿将其地理分布范围扩大到内蒙古。真角鹿是从非脱换型鹿角向季节性脱换型鹿角过渡的代表，在欧亚大陆都有分布，化德是继通古尔和四子王旗之后在内蒙古发现这个属的第三个地区。化德祖鹿的鹿角具有4个枝，远端两个枝剑形，在土城子地点是第二次发现，但目前尚未在其他地区发现过，似乎是化德一带的地方种类。山西祖鹿以其下臼齿不具古鹿褶而与新罗斯祖鹿(Cervavitus novorossiae)相区别。山西祖鹿在中国北方的分布较广，主要见于山西、陕西、甘肃和内蒙古，而且在每个地点的化石标本较多，指示其种群密度较大。布氏始柱角鹿、三角始柱角鹿和山西祖鹿是榆社盆地中榆社I带的主要鹿科成员，出现在岩石地层的马会组或生物年代学的保德期。真角鹿的时代分布主要在晚中新世。因此根据土城子地点的鹿科化石判断其地质时代应为晚中新世，而较多的山西祖鹿标本指示化德一带在晚中新世有较广的森林。
分类： 地球科学 >> 地质学 提交时间： 2017-08-08 合作期刊: 《古脊椎动物学报》
摘要： New specimens from a new locality at Chutoulang in eastern Nei Mongol were identifed as Canis chihliensis, Coelodonta nihowanensis, Hipparion (Proboscidipparion) sinense,Equus sanmeniensis, Sus lydekkeri, Muntiacus cf. M. lacustris, Axis shansius, Eucladoceros boulei, Spirocerus cf. S. wongi and Bison palaeosinensis. They enriched the mammalian fauna of Chutoulang to 30 taxa together with the material from Dongliang, Dongcun Beigou and Dongcun Nangou localities. The mammalian fossils from these four localities are all from the same horizon and can be regarded as the same fauna, i.e. Chutoulang fauna. The composition of Chutoulang fauna is the closest to that of Nihewan (s.s.) by binary faunal similarity coeffcients. The age of Chutoulang fauna is between those of Nihewan (s.s.) and Juyuandong at Liucheng according to Brainerd-Robinson’s sequence, extinction rates and antiquity coeffcients. The numerical age of Chutoulang fauna is estimated between 1.4–1.6 Ma based on those of compared faunas. Carnivora are the most numerous in Chutoulang fauna with 11 taxa, but mostly the small sized ones.Perissodactyla and Artiodactyla make about half of the fauna. They are mostly large sized forms. The presence of numerous browsers or forest dwellers implies the existence of forest or woodland in Chutoulang area during that period. The presence of grazers and openland dwellers indicates the existence of larger area of grassland or steppes than that of woodland or forests. Most members of Chutoulang fauna are temperate habitat dwellers with a few cold-prone forms such as Ochotona and Coelodonta. The climate in Chutoulang area in the Early Pleistocene was thus similar to that of today. Chutoulang fauna is the most northeastern Early Pleistocene fauna in China and it can be recommended as a type site of the Early Pleistocene mammalian fauna in northeastern China.
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